I don't have a technical reference for this, but I believe those are vents to prevent the cabin floor buckling in the event of a sudden pressurization failure.

The cargo hold is beneath the passenger cabin, and both are pressurized because the cylindrical hull can resist pressure better than the flat cabin floor. If the cabin suddenly loses pressure the air in the hold could buckle the floor upward, or if the hold loses pressure the air in the cabin could buckle the floor downward. The floor must be generously vented to prevent this.