Photos source: screenshot from youtube video ->
I think they can be used as fuselage reference points in flight testing, but I'm not sure.
Are they part of NATO standard symbology?
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It may or may not be being used in this application, but they are location markers, usually used in weapons separation tests to record the movement and behavior of stores in the airstream as they separate from the jet. Camera pods, usually mounted on the wingtips or other optimal vantage points record the process for engineering purposes.